Here are some variations:

left: leaf as demonstrated above.
center: leaf as demonstrated above except the shape is not pinched together in step 18. This allows the leaf to be round and the internal loops are more visible. The center mid-vein was cut off and the paper glued down inside the circle.
right: leaf made as above except in step 17, the shaped is folded so the loops face outwards. In order for the shape to maintain its shape, you need to wrap paper around the entire structure. Here, the wrap-around paper is blue
http://www.origami-resource-center.com/quilling-combing-leaf.html

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